[Pollinator] Pollen May Be Annoying, But It Is Extremely Important
Ladadams at aol.com
Ladadams at aol.com
Mon Jul 27 14:35:10 PDT 2009
from ThePilot.com
JUDY JESSOP: Pollen May Be Annoying, But It Is Extremely Important
Lightweight floating pollen that puffs with each step, speckling the air
we breathe, causes havoc with those of us that have allergies.
It is, however, the only way that many trees and plants achieve
pollination. Wind pollinators must dump huge amounts of pollen into the air.
This strategy is necessary to raise the possibility of some grains finding
the right flowers, so germination can take place. Ragweed is at the top of
the list of allergy, producing wind pollens. Others such as oak, birch,
maples and grasses may also trigger allergies. Now that summer has arrived,
though, and many wind-pollinators are finished, allergy symptoms are settling
down.
Not all pollen triggers hay fever. In general, those of us with such
allergies can be friends with all the plants that depend on insects, birds and
other animals to pollinate their flowers. Such plants have heavy, sticky
pollen that clings to any volunteers drawn to sip sweet nectar. The major ways
that pollen is transported in our region of the state is by insects, birds
and the wind.
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Flowers that depend on insects use many different strategies to attract
them. Some flowers generalize -- any insect will do, so their structure may
be quite simple, with no specialized features.
The tulip is a good example. It has a large wide cup of petals with long
anthers holding the pollen. Any insect crawling over the anthers, as it looks
for a sweet reward, will pick up a bit of sticky pollen to transport
onward.
Some flowers specialize to attract specific insects, such as bees. These
flowers often have a lobe that acts as a landing pad and deep-throated
petals. The pollen-laden anthers are located at the top of the flower throat,
dusting the backs of the bees as they enter. Bees are most sensitive to the
violet end of the color spectrum, so many bee-pollinated flowers have blue or
violet markings.
Another strategy, used by some flowers, is to display showy brackets, or
modified leaves, that attract attention (such as the dogwood). These bright
white flags also provide landing platforms for insects as they sip nectar
while pollinating the tiny flowers nestled in the center.
Flowers can develop some interesting relationships with their pollinators.
The plants of the yucca family, for instance, depend on a single species of
moth for successful pollination.
The female yucca moth gathers a ball of pollen and stuffs it into the
cup-shaped stigma of each flower. She then lays a few of her eggs in the pollen.
When the moth eggs hatch, the larva consume a small number of the
developing seeds, but this loss is outweighed by the benefit of having such an
effective pollinator.
Another interesting aspect of this symbiotic relationship is that if the
moth lays too many of her eggs on the stigma, the flower will abort and drop
off to the ground -- the plant's way of selecting against moths that
over-lay to the detriment of the yucca's regeneration.
Bird Pollination
Many flowers depend on hummingbirds for pollination. These flowers are
often deep throated, and the hummingbird hovers while reaching deep inside for
nectar.
Bird-pollinated flowers have no landing platform, since hummers need not
land to feed. Anthers are often dangled to dust the bird's head as he or she
feeds on the nectar. These flowers are often red, orange or deep pink,
colors that attract the birds but are inconspicuous to most insects.
The nectar of hummingbird flowers is relatively weak, about 25 percent
sugars with high levels of sucrose. Insect-pollinated flowers normally produce
highly concentrated nectars dominated by fructose and glucose.
Wind Pollination
As I mentioned earlier, many of the trees and plants that pollinate in
early spring depend on the wind to spread their pollen. These trees and plants
may be either gymnosperms (nonflowering) or angiosperms (flower-producing).
Pine trees are an excellent example of gymnosperms. These trees do not have
flowers. Instead, they reproduce by developing cones, which are modified
leaves. Some of these cones are male, producing pollen, while others are
female, with each scale of the cone housing two eggs.
The male cones are present only in early spring and now litter the ground.
Pine trees are particularly conspicuous when pollinating because the pollen
is yellow, and here in the Sandhills, we have lots and lots of pine trees
-- the reason that at times even the air swirls in yellow puffs.
Interestingly, pine pollen is not allergy-producing to most people. It just
gets blamed because there is so much and it is so yellow.
Other wind pollinators, the angiosperms, produce flowers, but the flowers
are not scented, and they don't produce nectar. The flowers are also very
simple and often do not even have petals. The red maple is an example. Often
these plants bloom before the leaves emerge to allow wind-borne pollen easy
access to the flowers.
The Precious Cargo
Pollen is amazing stuff, each grain a complicated little travel capsule.
Its important occupants are two male sperm cells.
The outside wall (called the exine) is very tough material composed of
waxes and proteins. It protects its precious cargo from solar radiation and
dehydration, as the cell bumps around in search of the right flower for
pollination. Exines have either pores or special ridges where the case is
thinner -- an important feature when the pollen finds the right mate.
Packed inside the exine is an inner layer of cellulose (the intine) much
like a cell wall. Inside this layer are a few cells. One of these cells is
very specialized. It is a reproductive cell with two nuclei. One nucleus will
produce a pollen tube, and one will divide into two sperm cells.
In angiosperms, every grain of pollen that successfully reaches the flower
of its own species still has work to do. When it arrives on the stigma (the
part of a flower that receives pollen), the tube cell in this grain of
pollen begins to elongate, exiting the outer wall of the pollen grain through
a pore or ridge.
The tube continues growing down into the center of the flower toward the
unfertilized eggs. This single-celled tube can be quite long in some plants.
In corn, for example, the tube must grow as much as 12 inches to reach the
eggs.
As the tube grows, the two sperm cells travel down the tube toward the tip.
When the tip of the tube reaches an ovule (female reproductive cell), it
bursts, releasing the two sperm cells.
One of these sperm cells will unite with the egg, making a fertile seed.
The other will unite with another cell, which produces food that is stored
around the seed. The food store is high in starch, protein and oils. Such
grasses as corn are so high in stored food that they have become a main staple
of the human diet.
In gymnosperms (such as our pines), during the period of time that pollen
is being released to the wind, the scales of female cones open temporarily
to receive pollen, and the pollen grain is drawn directly into the egg. The
female cones then close during fertilization and maturation.
There are many intricate, amazing and important things that are going on
during pollination -- each single microscopic pollen grain a complex
structure. Yet one pinch of pollen powder holds thousands of grains. It may at
times wreak havoc with those of us with allergies, but without it where would
we be?
Contact Judy Jessop by e-mail at jgjessop at gmail.com.
Laurie Davies Adams
Executive Director
Pollinator Partnership
423 Washington Street, 5th floor
San Francisco, CA 94111
415-362-1137
LDA at pollinator.org
_www.pollinator.org_ (http://www.pollinator.org/)
_www.nappc.org_ (http://www.nappc.org/)
National Pollinator Week is June 22-28, 2009.
Beecome involved at _www.pollinator.org_ (http://www.pollinator.org/)
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